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Danny’s DNA Discoveries – Tricholomatineae Incertae Cedis of the PNW
by Danny Miller

Introduction

These are in uncertain families of the suborder Tricholomatineae. They may each get their own families (seems likely), or they may be placed in an existing family. Nothing much unites these yet morphologically.

TBD - everything except Pseudoomphalina angelesiana and 'Cleistocybe' gomphidioides.

abundant common uncommon rare - colour codes match my Pictorial Key and are my opinions and probably reflect my bias of living in W WA. Rare species may be locally common in certain places at certain times.

Click here to download the FASTA data of all my DNA sequences

Summary of Interesting Results
  • tbd

Aspropaxillus - click to expand

One of our largest gilled mushrooms, a huge funnel-shaped clitocyboid up to 40cm across. Clitocybe maxima and Infundibulicybe geotropa may also get large. Unpleasant odor and taste. Formerly in Leucopaxillus because the gills separate easily, but although large, these are not as dense as Leucopaxillus. Amyloid spores.

Species mentioned: Aspropaxillus giganteus, septentrionalis.

Cleistocybe gomphidioides - click to expand

'Cleistocybe' gomphidioides WA - this clitocyboid with a veil can be told apart from Catathelasma and true Cleistocybe by viscid cap, growth in fall, and a sometimes fleeting veil/ring, making it very hard to ID if you don't notice it. Strongly rancid-farinaceous. True Cleistocybe, such as Cleistocybe vernalis have a dry cap. This species needs to be moved into a new genus and family.

Omphaliaster - click to expand

Recognized by round spores with warty nodules, but otherwise a nondescript omphalinoid/clitocyboid with an indented hygrophanous cap that may be striate, somewhat decurrent gills and somewhat farinaceous odor and taste.

Species mentioned: Omphaliaster asterosporus, borealis.

Paralepista - click to expand

Orange, strongly funnel-shaped clitocyboids with close/crowded gills and almost round spores that are off-white and finely warty (like Lepista, the name given to some Clitocybe species with similarly coloured warty spores). Bonomyces is stockier and smells farinaceous and has smooth, elliptical spores. Infundibulicybe gibba is pinkish tan but not as orange, and has smooth, white spores.

Species mentioned: Paralepista flaccida, gilva, inversa

Pseudoomphalina - click to expand

Not only are they hard to tell from other genera (and even mushrooms in other families) they don't look anything alike themselves. They were separated from Clitocybe by having amyloid spores and cystidia along with the similar Pseudolaccaria, but some European authors say both segregate genera lack cystidia so at best the cystidia are hard to find and there are no good separating characters besides this being one of the many genera that, unlike Clitocybe, have amyloid spores.

Species mentioned: Pseudoomphalina angelesiana, intermedia, kalchbrenneri. Neohygrophorus angelesianus.

Rimbachia - click to expand

Stemless oysters on moss with rudimentary or no gills, usually white. They are difficult to distinguish from some Arrhenia species (usually not white) and also Muscinupta (white with a tapered fluting "stem"). It's possible they belong in the Omphalinaceae, but we'll see.

Species mentioned: Rimbachia arachnoidea, bryophila, neckerae

Ripartites - click to expand

A beautiful clitocyboid with a white hairy cap with a bearded, inrolled margin, strongly decurrent gills that turn pinkish brown in age from pale brown spores and a rose-tinted powdery stem. Note especially that this is not white spored!

Species mentioned: Ripartites tricholoma.

 

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